Anubias
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{{Taxobox
| name = ''Anubias''
| image = Anubias_barteri_var_nana_on_a_bogwood.jpg
| image_width = 240px
| image_caption = ''Anubias barteri'' var. ''nana'' on a bogwood (top view)
| regnum = [[Plant]]ae
| divisio = [[Flowering plant|Magnoliophyta]]
| classis = [[Liliopsida]]
| ordo = [[Alismatales]]
| familia = [[Araceae]]
| genus = '''''Anubias'''''
| genus_authority = [[Heinrich Wilhelm Schott|Schott]]
| subdivision_ranks = Species
| subdivision =
*''[[#Anubias afzelii|Anubias afzelii]]''
*''[[#Anubias barteri|Anubias barteri]]''
**''Anubias barteri'' var. ''angustifolia''
**''Anubias barteri'' var. ''barteri''
**''Anubias barteri'' var. ''caladiifolia''
**''Anubias barteri'' var. ''glabra''
**''Anubias barteri'' var. ''nana''
*''[[#Anubias gigantea|Anubias gigantea]]''
*''[[#Anubias gilletii|Anubias gilletii]]''
*''[[#Anubias gracilis|Anubias gracilis]]''
*''[[#Anubias hastifolia|Anubias hastifolia]]''
*''[[#Anubias heterophylla|Anubias heterophylla]]''
*''[[#Anubias pynaertii|Anubias pynaertii]]''
}}
'''''Anubias''''' is a genus of aquatic and semi-aquatic [[flowering plant]]s in the family [[Araceae]], native to tropical central and western [[Africa]]. They primarily grow in rivers and streams, but can also be found in marshes. They are characterized by broad, thick, dark leaves that come in many different forms. The genus was revised in 1979<ref name="Mededelingen">W. Crusio (1979) A revision of Anubias Schott (Araceae). ''Mededelingen Landbouwhogeschool Wageningen'' '''79''' (14) 1-48</ref> and since then its [[taxonomy]] has been stable. Species can be determined by using mostly characteristics of the [[inflorescence]].
=== Reproduction and uses ===
''Anubias'' are commonly used in [[aquarium]]s, usually attached to rocks or bogwood. In contrast to most plants, ''Anubias'' generally prefer subdued lighting and can also produce flowers underwater. In the aquarium they should be placed in shaded areas, otherwise algae will develop on the leaves.
''Anubias'' are considered by many aquarists some of the easiest plants to maintain, since their light and nutrient requirements are very low and also because [[herbivore|herbivorous]] fish will not eat it. This is why ''Anubias'' are some of the few plants which can be used in aquariums with African [[cichlid]]s and [[goldfish]]es.
Reproduction in artificial environments can be accomplished by stolon division or from side shoots. The [[stolon]] must always be above the substrate in order to survive, otherwise it will rot and the plant dies. It is also possible to propagate ''Anubias'' by seed<ref name="Anubias">W. Crusio (1987) Die Gattung Anubias SCHOTT (Araceae). ''Aqua-Planta'' '''Sonderheft 1''' 1-44</ref>.
The natural growth rate of all species in this genus is rather slow. Usually, they produce a leaf every 3 weeks, or even slower. Unfortunately, these are also among the few plants that do not respond to [[Aquarium CO2 addition|CO<sub>2</sub> addition]].
The most commonly available species of this genus is ''Anubias barteri'' Schott, which is very polymorphic and is subdivided into several varieties. The biggest representatives of the genus are ''Anubias gigantea'' Chevalier ex Hutchinson and ''Anubias heterophylla'' Engler. Their leaf-stems can grow up to 83 cm, with leaves 40 cm long and 14 cm broad with lateral lobes up to 28 cm long and 10 cm broad. The smallest representative is ''[[Anubias barteri var. nana]]'' (Engler) Crusio, with a height up to 10 cm and with leaves up to 6 cm long and 3 cm broad.
''Anubias'' can best be grown emersed (above water). For this reason they may be used in [[Vivarium|paludarium]]s.
== Quick facts ==
;''Anubias afzelii'' Schott
* Narrow-leafed, medium-sized Anubias.
* Characteristics: plant with stolon.
* Leaf-stem: uo to 20 cm
* Leaves: 13-35 cm long, 3-13 cm broad
* Height in aquarium: 25 - 30 cm
Optimal conditions:
* Temperature: 22 - 27 °C
* pH: 6.0 - 7.0
* Water hardness: 2 dGH - 6 dGH
* Position in aquarium: middle to back
* Usual growth rate: one leaf every 2 months
;''Anubias barteri'' Schott
Common varieties:
;''Anubias barteri'' var. ''angustifolia'' (Engler) Crusio
* Synonym: ''Anubias lanceolata'' f. ''angustifolia'' Engler.
* Narrow leafs, similar to ''afzelii'', but much smaller
* Leaf-stem: up to 32 cm
* Leaves: up to 18 cm long, up to 3.5 cm wide
* Height in aquarium: 10 - 15 cm
Optimal conditions:
* Temperature: 20 - 27 °C
* pH: 5.5 - 8.0
* Water hardness: < 8 dGH
* Position in aquarium: middle
* Usual growth rate: one leaf every 2 months
;[[Anubias barteri var. barteri|''Anubias barteri'' var. ''barteri'']]
* Compact, heart-shaped anubias
* Leaf-stem: up to 23 cm
* Leaves: 7-23 cm long, 4-11 cm wide
* Height in aquarium: 25 - 45 cm
Optimal conditions:
* Temperature: 20 - 27 °C
* pH: 5.5 - 9.0
* Water hardness: < 20 dGH
* Position in aquarium: back
* Usual growth rate: one leaf every 3 months
;''Anubias barteri'' var. ''caladiifolia'' Engler
* Heart-shaped anubias
* Leaf-stem: up to 54 cm
* Leaves: 10-23 cm long, 5-14 cm wide
* Height in aquarium: 7 - 30 cm
Optimal conditions:
* Temperature: 20 - 27 °C
* pH: 5.5 - 8.0
* Water hardness: < 20 dGH
* Position in aquarium: middle to back
* Usual growth rate: one leaf every 2 months
;''"Anubias barteri'' var. ''coffeefolia"''
* New leaves are red-brown
Several varieties of ''Anubias barteri'' (and also some specimens of some other species) will sometimes produce red-brown leaves. Although these plants have been traded under the name ''"Anubias barteri'' var. ''coffeefolia"'', this name has no taxonomic status. Leave color is not important in determining to which ''Anubias'' species a certain plant belongs.<ref name="Mededelingen"/>
;''Anubias barteri'' var. ''glabra'' N. E. Brown
* Synonyms: ''Anubias lanceolata'' N. E. Brown, ''Anubias minima'' Chevalier.
* Narrow-leafed, large anubias
* Leaf-stem: up to 35 cm long
* Leaves: spear-shaped, up to 21 cm long, 9 cm wide
* Height in aquarium: 30 - 50 cm
Optimal conditions:
* Temperature: 22 - 27 °C
* pH: 5.5 - 8.0
* Water hardness: < 20 dGH
* Position in aquarium: back
* Usual growth rate: 4 - 8 leaves per year
;[[Anubias barteri var. nana|''Anubias barteri'' var. ''nana'' (Engler) Crusio]]
* Synonym: ''Anubias nana'' Crusio
* Dwarf, creeping, with heart-shaped leaves
* Leaf-stem: up to 5 cm long
* Leaves: up to 6 cm long and 3 cm wide
* Height in aquarium: 5 - 10 cm
Optimal conditions:
* Temperature: 22 - 27 °C
* pH: 5.5 - 9.0
* Water hardness: 3 - 10 dGH
* Position in aquarium: front
* Usual growth rate: one leaf every month
;''Anubias gigantea'' Chevalier ex Hutchinson
* Synonyms: ''Anubias gigantea'' var. ''tripartita'' Chevalier, ''Anubias hastifolia'' var. ''robusta'' Engler
* Large arrow-shaped leaves
* Leaf-stem: up to 83 cm long
* Leaves: up to 30 cm long and 14 cm wide, with large lateral lobes up to 28 cm long and 10 cm broad
* Seldomly used in aquariums
;''Anubias gilletii'' De Wildeman & Durand
* Initially heart-shaped, later with long rear fringes.
* Leaf-stem: up to 40 cm
* Leaves: arrow-shaped, 25 cm long, 12 - 13 cm wide, lateral lobes up to 13 cm long
* Height in aquarium: 25 - 40 cm
Optimal conditions:
* Temperature: 22 - 27 °C
* pH: 6.0 - 8.0
* Water hardness: 4 - 10 dGH
* Position in aquarium: back
* Usual growth rate: 2 - 6 leaves per year
;[[Anubias gracilis|''Anubias gracilis'' Chevalier ex Hutchinson]]
* Whether this is a separate species or another variety of ''Anubias barteri'' is doubtful<ref name="Anubias" />
* Leaf-stem: up to 33 cm
* Leaves: triangular heart-shaped, 12 cm long, 4-10 cm wide, lateral lobes up to 7 cm long and 3 cm wide
* Height in aquarium: 20 - 30 cm
Optimal conditions:
* Temperature: 24 - 27 °C
* pH: 6.0 - 8.0
* Water hardness: 5 - 12 dGH
* Position in aquarium: middle to back
* Usual growth rate: 2 - 6 leaves per year
;''Anubias hastifolia'' Engler
* Synonyms: ''Amauriella hastifolia'' (Engler) Hepper, ''Anubias hastifolia'' var. ''sublobata'' Engler, ''Anubias auriculata'' Engler, ''Amauriella auriculata'' (Engler) Hepper, ''Anubias haullevilleana'' De Wildeman, ''Anubias laurentii'' De Wildeman, ''Amauriella obanensis'' Rendle, ''Amauriella talbotii'' Rendle
* Leaves: long heart-shaped
* Leaf-stem: up to 67 cm long
* Leaves: up to 33 cm long and 14 cm wide, lateral lobes up to 26 cm long and 8 cm broad
* Height in aquarium: 30 - 50 cm
Optimal conditions:
* Temperature: 22 - 27 °C
* pH: 6.0 - 8.0
* Water hardness: < 20 dGH
* Position in aquarium: middle to back
* Usual growth rate: 2 - 6 leaves per year
[[Image:Anubias heterophylla..JPG|thumb|200px|right|''Anubias heterophylla'']]
;''Anubias heterophylla'' Engler
* Synonyms: ''Anubias congensis'' N. E. Brown, ''Anubias congensis'' var. ''crassispadix'' Engler, ''Anubias affinis'' De Wildeman, ''Anubias engleri'' De Wildeman, ''Anubias bequertii'' De Wildeman, ''Anubias undulata'' (trade name)
* Very large species, for tall aquariums
* Leaf-stem: up to 66 cm long
* Leaves: 38 cm long, 13 cm wide, sometimes with very short basal lobes
* Height in aquarium: 25 - 60 cm
Optimal conditions:
* Temperature: 24 - 27 °C
* pH: 5.5 - 8.0
* Water hardness: 5 - 12 dGH
* Position in aquarium: back
* Usual growth rate: 2 - 4 leaves per year
;''Anubias pynaertii'' De Wildeman
* Leaf-stem: up to 45 cm
* Leaves: up to 29 cm long and 14 cm broad
== See also ==
* [[Aquatic plant]]
*[[List of freshwater aquarium plant species]]
==References==
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[[Category:Araceae]]
[[Category:Aquatic plants]]
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