Badamia exclamationis
3698910
221062644
2008-06-22T22:07:09Z
Dysmorodrepanis
303688
/* References */ restore alphabetic refs (author/date are more useful than publishing medium)
{{Taxobox | name = Brown Awl
| status =
| image = Brown awl.jpg
| image_width = 240px
| image_caption =
| regnum = [[Animal]]ia
| phylum = [[Arthropod]]a
| classis = [[Insect]]a
| ordo = [[Lepidoptera]]
| familia = [[Hesperiidae]]
| subfamilia = [[Coeliadinae]]
| genus = ''[[Badamia]]''
| species = '''''B. exclamationis'''''
| binomial = ''Badamia exclamationis''
| binomial_authority = ([[Johan Christian Fabricius|Fabricius]], 1775)<ref name="LepIndex Badamia exclamationis">Card for [http://internt.nhm.ac.uk/jdsml/research-curation/projects/lepindex/detail.dsml?TaxonNo=181283.0&UserID=&UserName=&&listPageURL=list%2edsml%3fsort%3dSCIENTIFIC%255fNAME%255fon%255fcard%26SCIENTIFIC%5fNAME%5fon%5fcardqtype%3dstarts%2bwith%26SCIENTIFIC%5fNAME%5fon%5fcard%3dexclamationis%26recLimit%3d30&searchPageURL=index%2edsml%3fSCIENTIFIC%5fNAME%5fon%5fcardqtype%3dstarts%2bwith%26sort%3dSCIENTIFIC%255fNAME%255fon%255fcard%26SCIENTIFIC%5fNAME%5fon%5fcard%3dexclamationis%26recLimit%3d30 '''''Badamia exclamationis'''''] in LepIndex. Accessed 12 October 2007.</ref>
| synonyms =
| range_map =
| range_map_width = 250px
| range_map_caption =
}}
'''''Badamia exclamationis''''', commonly known as the '''Brown Awl''',<ref name="Savela Badamia exclamationis">Marrku Savela's Website on Lepidoptera. Page on genus [http://www.funet.fi/pub/sci/bio/life/insecta/lepidoptera/ditrysia/hesperioidea/hesperiidae/coeliadinae/badamia/index.html ''Badamia''].</ref><ref name="TOLWeb Badamia">TOL web page on genus [http://www.tolweb.org/Badamia/94258 ''Badamia'']</ref> is a [[butterfly]] belonging to the family [[Hesperiidae]], which is found in [[India]], [[Southeast Asia]] and [[Australia]].
==Range==
The Brown Awl is found in [[Sri Lanka]], [[India]], [[Myanmar]], South [[Yunnan]], [[Australia]] and [[Japan]].<ref name="Savela Badamia exclamationis"/>
The butterfly ranges over the complete [[Indian subcontinent]] and is also found in the [[Andaman islands]].<ref name="Savela Badamia exclamationis"/><ref name="Evans">Evans,W.H.(1932) The Identification of Indian Butterflies, ser no I 5.1, pg 321.</ref><ref name="Kunte">Kunte, Krushnamegh. (2000) Butterflies of Peninsular India, ser no 63, pp 194-196.</ref>
The type locality is [[South India]].<ref name="Savela Badamia exclamationis"/>
==Status==
As per Evans (1932), the butterfly is 'Common' in [[India]] and 'Rare' in the [[Andaman islands]].<ref name="Evans"/> Wynter-Blyth (1957) records it as 'Not Rare' and 'Locally Common'.<ref name="Wynter-Blyth">Wynter-Blyth, M.A. (1957) Butterflies of the Indian Region, pg 470-471.</ref> Kunte (2000) reports it as common in deciduous forests during the monsoon months and the evergreen forests in the following months.<ref name="Kunte"/>
==Habits==
A forest butterfly, the Brown Awl favours openings and edges of deciduous and evergreen forests while its caterpillars are to be found in moist deciduous and semi-evergreen forests.<ref name="Kunte"/> It flies about either late or early in the morning in the shade of the jungles.<ref name="Wynter-Blyth"/> It can be sometimes seen in bright sunlight visiting flowers<ref name="Wynter-Blyth"/>, such as ''Glycosmis'' ''Buddleia'', ''Chromolaena'' and ''Lantana'', but is very wary and energetic at such times, moving jerkily and rapidly between flowers or across infloresences. It can also be seen mud-puddling or at bird-droppings.<ref name="Kunte"/>
During a population explosion, like those of the Common Banded Awl, the caterpillars of the Brown Awl may strip away all their food supply forcing the butterflies to migrate to other places where fresh supply of host-plants are available and even into other habitats such as shrubs, grasslands and gardens.<ref name="Kunte"/>
The Brown Awl flies as low as 6 feet over the bushes or as high as 60 to 75 feet in the canopy. The adults feed at lower levels on flowers of shrubs and small trees, but ascend to higher reaches of the vegetation to lay eggs or to bask, which it does very occasionally, holding its wings flat with the forewings covering the hindwngs thus giving an arrowhead effect. The flight of the butterfly is fast and bounding with an audible wing-beat.<ref name="Kunte"/>
When inactive, it rests on the undersides of leaves in shady forest spots, with the head pointing downwards. If disturbed it will generally buzz around energetically befor returning to the same spot to rest.<ref name="Kunte"/>
==Description==
:''See [[glossary of Lepidopteran terms|glossary]] for terms used''
The Brown Awl is a non-descript brown butterfly, darker above and lighter below. The sexes are alike, except for 3 to 4 semi-transparent spots on the forewing which cannot be differentiated in the field. The skipper has a light-brown abdomen with black bands across it. The DSF is usually smaller, paler, and may not have the forewing spots.<ref name="Kunte"/>
This skipper is unmistakable because of its long and narrow wings. It has the longest wings in proportion to breadth of all Indian butterflies.<ref name="Kunte"/>
====Detailed description====
Watson (1891) gives a detailed description:<ref name=watson>Watson, E. Y. (1891) Hesperiidae indicae.</ref>
'''Genus characters'''
:''Forewing, narrow, elongated ; costa slightly arched at base, exterior margin very oblique and slightly convex below the apex ; cell very long and narrow, extending three-fourths the wing ; first subcostal branch emitted at two-fifths, second at one-fifth, third at one-seventh, fourth close to and fifth at end of the cell ; disco-cellulars very slender, inwardly oblique, of nearly equal length, upper bent inward close to subcostal ; upper radial from the angle, lower from their middle ; median branches curved at their base, middle branch emitted at about one-fourth, and lower at three-fourths before end of the cell ; submedian curved in the middle ; hindwing short ; apex very convex, angularly lobed at anal angle, abdominal margin short ; precostal projecting inward ; costal vein arched upward from the base ; second subcostal emitted at one-third from the base ; cell broad throughout ; disco-cellulars very slender, scarcely visible, of equal length ; radial from their angle, very slender ; middle median at about one-third, and lower at one-fifth from the base ; submedian straight, internal slightly curved. Thorax stout ; abdomen rather long, attenuated ; head broad ; palpi broad and flattened in front, bristly on outer edge, third joint long,projected forward, cylindrical ; fore-tibiae tufted beneath, femora slightly pilose beneath ; antennae with a lengthened club and long pointed tip.''
'''Species description'''
:"Upperside dark purplish brown, the base of both wings greyish olive brown.''
:'''Male'''. ''Forewing with three transparent slender yellow spots disposed longitudinally on the upper disc, the inner spot ending within the cell.''
:'''Female'''. ''Forewing with the spots larger, the middle spot oblique and irregularly angulated ; a less distinct spot also above the middle of sub-median vein. Underside pale greyish brown : forewing with discal area darker brown, the spots as above, and pale ochreous posterior border: hindwing with a dark brown anal area bordered above by a short pale ochreous streak. Thorax greyish olive brown ; abdomen dark brown with pale ochreous segmental bands; head and palpi in front pale ochreous with brown streaks; third joint of palpi brown ; legs brown above, pale beneath."
==Similar Species==
Unlike the other Awls, the Brown Awl lacks the narrow white wing bands on the hind wings. The very distinctive characteristics of the Brown Awl are the characteristic shape of the body and the narrower wings than the other Awls.<ref name=Kunte/>
==Life cycle==
====Eggs====
The Brown Awl lays many eggs on a single plant, one at a time, on the tips of fresh shoots. The dome-shaped egg is pale green with longitudinal ridges having fine beadings; a total 13 ridges in all.
====Larva====
The larva is a pale violaceous yellow, with numerous black transverse dorsal lines ; the prolegs are whitish encircled with black.<ref name=watson/> The head is yellow, approximately heart-shaped, with a black band and many tiny black spots.<ref name=Kunte/>
On hatching the larva webs the edges of leaves together with silk<ref name="Haribal">Haribal, Meena (1992) Butterflies of Sikkim, ser 541, pg 195-196.</ref> to form a roomy cell from a leaf in which it resides throughout the larval stage. When disturbed, it can quite briskly and even drop off. The caterpillars of the Brown Awl grow faster than most of those of other families, and have moist, sticky droppings.<ref name=Kunte/>
At the time of pupation they descend close to the ground, looking for suitable spots to pupate. The caterpillar constructs a tubular cell from a leaf by drawing the edges together with thick strands of silk.<ref name="Haribal"/> In this cell, the caterpillar prepares an extensive silk bed on which it sits awaiting pupation. The freshly formed pupa clings onto the silken pad almost immediately.<ref name=Kunte/>
====Pupa====
The Pupa is stubby, with protruding eyes and a prominent projection on the head in between them.<ref name=Kunte/> The pupa may be light-brown<ref name=Kunte/> or violaceous<ref name=watson/> in colour. The body tapers away from the shoulders towards the rear. The abdomwn is creamish with a row of four black spots on each side. The pupa is shiny, but plastered with a white powder.<ref name=Kunte/>
====Hostplants====
The caterpillars have been recorded on the following deciduous and semi-evergreen forest plants, mostly from the family ''[[Combretaceae]]''<ref name=Kunte/>:
* Large climbing shrubs of ''[[Combretum]]'' such as ''[[Combretum albidum]]''<ref name=Kunte/>, ''[[Combretum latifolium]]''<ref name=Kunte/> and ''[[Combretum ovalifolium]]''<ref name="Savela Badamia exclamationis"/>.
* Large forest trees of ''[[Terminalia]]'' genus such as ''[[Terminalia bellerica]]''<ref name=Kunte/><ref name="Savela Badamia exclamationis"/>, ''[[Terminalia catappa]]''<ref name="Savela Badamia exclamationis"/> and ''[[Terminalia oblongata]]''<ref name="Savela Badamia exclamationis"/>.
* ''[[Linociera purpurea]]''.<ref name="Savela Badamia exclamationis"/>
* ''[[Ficus]]'' spp.<ref name="Savela Badamia exclamationis"/>
== Gallery==
<gallery>
image:Badamia exclamationis (ento-csiro-au).jpg|Brown Awl
Image:ExclamationisCRW 5480-01.jpg|Museum specimens
</gallery>
==Cited references==
{{reflist}}
==See also==
*[[Hesperiidae]]
*[[Coeliadinae]]
*[[List of butterflies of India (Coeliadinae)]]
*[[List of butterflies of India (Hesperiidae)]]
==References==
====Print====
*{{aut|Beccaloni, G. W.}}, {{aut|Scoble, M. J.}}, {{aut|Robinson, G. S.}} & {{aut|Pitkin, B.}} (Editors). 2003. The Global Lepidoptera Names Index (LepIndex). World Wide Web electronic publication. [http://www.nhm.ac.uk/entomology/lepindex] (accessed 22 September 2007).
*{{aut|Brower, Andrew V. Z.}} and {{aut|Warren, Andrew}}, (2007). Coeliadinae Evans 1937. Version 21 February 2007 (temporary). http://tolweb.org/Coeliadinae/12150/2007.02.21 in The Tree of Life Web Project, http://tolweb.org/
*{{aut|Evans, W.H.}} (1932) ''The Identification of Indian Butterflies''. 2nd Ed, (i to x, pp454, Plates I to XXXII), Bombay Natural History Society, Mumbai, India.
*{{aut|Kunte, Krushnamegh}}. (2000) ''Butterflies of Peninsular India'', (i to xviii, pp254, Plates 1 to 32) Universities Press (India) Ltd, Hyderabad (reprint 2006). ISBN 81-7371-354-5.
*{{aut|Savela, Marrku }} Website on Lepidoptera [http://www.nic.funet.fi/pub/sci/bio/life/index.html] (accessed 22 September 2007)
*{{aut|Watson, E. Y.}} (1891) ''Hesperiidae indicae''. Vest and Co. Madras.
*{{aut|Wynter-Blyth, M.A.}} (1957) ''Butterflies of the Indian Region'', Bombay Natural History Society, Mumbai, India.
[[Category:Coeliadinae]]