Short-beaked Echidna
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{{Taxobox
| name = Short-beaked Echidna<ref name=msw3>{{MSW3 Groves|pages = 1-2|id=10300004}}</ref>
| status = LR/lc
| status_system = iucn2.3
| trend = stable
| status_ref = <ref>{{IUCN2006|assessors=Australasian Marsupial & Monotreme Specialist Group|year=1996|id=41312|title=Tachyglossus aculeatus|downloaded=12 May 2006}} Database entry includes justification for why this species is of least concern </ref>
| image = Echidna ST 03.jpg
| regnum = [[Animal]]ia
| phylum = [[Chordate|Chordata]]
| classis = [[Mammal]]ia
| subclassis = [[Prototheria]]
| ordo = [[Monotreme|Monotremata]]
| familia = [[Tachyglossidae]]
| genus = '''''Tachyglossus'''''
| genus_authority = [[Johann Karl Wilhelm Illiger|Illiger]], 1811
| species = '''''T. aculeatus'''''
| binomial = ''Tachyglossus aculeatus''
| binomial_authority = ([[George Shaw|Shaw]], 1792)
}}
The '''Short-beaked Echidna''' ('''''Tachyglossus aculeatus'''''), also known as the '''Spiny Anteater''' because of its diet of [[ant]]s and [[termites]], is one of four living species of [[echidna]] and the only member of the genus '''''Tachyglossus'''''. The Short-beaked Echidna is covered in fur and spines and has a distinctive snout and a specialized tongue, which it uses to catch its prey at a great speed. Like the other extant [[monotreme]]s, the Short-beaked Echidna lays [[egg (biology)|eggs]]; the monotremes are the only group of [[mammal]]s to do so.
The species is found throughout [[Australia]], where it is the most widespread native mammal, and in coastal and highland regions of southwestern [[New Guinea]], where it is known as the ''Mungwe'' in the [[Dadibi language|Daribi]] and [[Kuman language (New Guinea)|Chimbu]] languages.<ref name = "Flannery">Flannery, T. 1990. ''Mammals of New Guinea'' Robert Brown & Associates ISBN 1-86273-029-6</ref> It is not threatened with extinction, but human activities, such as hunting, habitat destruction, and the [[Invasive species in Australia|introduction]] of foreign predatory species and parasites, have reduced the distribution of the Short-beaked Echidna in Australia.
==Taxonomy and naming==
The Short-beaked Echidna was first described by [[George Shaw]] in 1792. He named the [[species]] ''Myrmecophaga aculeata'', thinking it might be related to the South American [[anteater]]. Since Shaw first described the species, its name has undergone four revisions: from ''M. aculeata'' to ''Ornithorhynchus hystrix'', ''Echidna hystrix'', ''Echidna aculeata'' and, finally, ''Tachyglossus aculeatus''.<ref>Iredale, T. and Troughton, E. 1934. A checklist of mammals recorded from Australia. ''Memoirs of the Australian Museum'' 6: i–xii, 1–122</ref> The name ''Tachyglossus'' means "quick tongue", in reference to the speed with which the Echidna uses its tongue to catch ants and termites, and ''aculeatus'' means "spiny" or "equipped with spines".
The Short-beaked Echidna is the only member of its [[genus]], sharing the family [[Echidna|Tachyglossidae]] with the extant species of the genus ''[[Zaglossus]]'' that occurs in New Guinea. ''Zaglossus'' species, which includes the [[Western Long-beaked Echidna|Western Long-beaked]], [[Sir David's Long-beaked Echidna|Sir David's Long-beaked]] and [[Eastern Long-beaked Echidna]], are all significantly larger than ''T. aculeatus'', and their diet consists mostly of worms and grubs rather than ants and termites. Species of the Tachyglossidae are egg-laying mammals; together with the related family [[Platypus|Ornithorhynchidae]], they are the only extant [[monotreme]]s in the world.
There are five subspecies of the Short-beaked Echidna, each found in a different geographical location. The subspecies also vary from each other in terms of hairiness, spine length and width, and the size of the grooming claws on their hind feet.
*'''''T. a. multiaculeatus''''' is found on [[Kangaroo Island]];
*'''''T. a. setosus''''' is found on [[Tasmania]] and some islands in [[Bass Strait]];
*'''''T. a. acanthion''''' is found in [[Northern Territory]] and [[Western Australia]];
*'''''T. a. aculeatus''''' is found in [[Queensland]], [[New South Wales]], [[South Australia]] and [[Victoria (Australia)|Victoria]];
*'''''T. a. lawesii''''' is found in coastal regions and the highlands of New Guinea, and possibly in the rainforests of Northeast Queensland.<ref>Griffiths, M.E. 1978. ''The Biology of Monotremes''. Academic Press : New York ISBN 0-12-303850-2</ref>
The Short-beaked Echidna was commonly called the Spiny Anteater in older books, though this term is has fallen out of fashion as it bears no relation to the true anteaters. It has a variety of names among the languages of the regions where it is found. The [[Noongar|Noongar people]] from southwestern [[Western Australia]] call it the ''Nyingarn''. In [[Central Australia]] southwest of [[Alice Springs]], the [[Pitjantjatjara language|Pitjantjatjara]] term is ''tjilkamata'' or ''tjirili'', from the word ''tjiri'' for spike of Porcupine Grass ''([[Triodia irritans]])''. The word can also mean slowpoke.<ref>{{cite book |author=Cliff Goddard|title= Pitjantjatjara/Yankunytjatjara To English Dictionary |year=1992|edition=2||pages=152|publisher=Institute for Aboriginal Development |location=Alice Springs, Northern Territory |isbn= 0-949659-64-9}}</ref> It is called ''(minha) kekoywa'', where ''minha'' is a qualifier meaning 'meat' or 'animal', in Pakanh, ''(inh -) ekorak'' in Uw Oykangand, and ''(inh -) egorag'' in Uw Olkola, where ''inh-'' is a qualifier meaning 'meat' or 'animal' three aboriginal languages of central [[Cape York Peninsula]].<ref name ="PH1997">{{cite web | author = Philip Hamilton | title = short-beaked echidna, ''Tachyglossus aculeatus'' | publisher = Australian Institute of Aboriginal and Torres Strait Islander Studies | date = 1997 | url = http://www.geocities.com/Athens/Delphi/2970/echidna.htm | accessdate = 2007-06-08}}</ref>
In the highland regions of southwestern [[New Guinea]] it is known as the ''Mungwe'' in the [[Dadibi language|Daribi]] and [[Kuman language (New Guinea)|Chimbu]] languages.<ref name = "Flannery">Flannery, T. 1990. ''Mammals of New Guinea'' Robert Brown & Associates ISBN 1-86273-029-6</ref>
==Description==
Short-beaked Echidnas are typically 30 to 45 centimetres (12-18 in) in length, have a 75-millimetre (3 in) snout, and weigh between two and five [[kilogram]]s (4.5-11 lb). However, the Tasmanian subspecies, ''T. a. setosus'', is larger than its Australian mainland counterparts. Because the neck is not externally visible, the head and body appear to merge together. The earholes are on either side of the head, with no external [[pinna (anatomy|pinnae]]. The eyes are small and at the base of the wedge-shaped snout. The nostrils and the mouth are at the distal end of the snout; the mouth of the Short-beaked Echidna cannot open wider than 5 mm.<ref name = "Murray''>Murray, P.F. 1981. A unique jaw mechanism in the echidna, ''Tachglossus aculeatus'' (Monotremata). ''Australian Journal of Zoology'' 29: 1–5</ref> The body of the Short-beaked Echidna is, with the exception of the underside, face and legs, covered with cream-coloured spines. The spines, which may be up to 50 mm (2 in) long, are modified hairs, mostly made of [[keratin]]. Insulation is provided by [[fur]] between the spines, which ranges in colour from honey to a dark reddish-brown and even black; the underside and short tail are also covered in fur. Colouration of the fur and spines varies with geographic location. The Echidna's fur may be infested with what is said to be the world's largest [[flea]], ''[[Echidna flea|Bradiopsylla echidnae]]'', which is about 4 mm long.
The limbs of the Short-beaked Echidna are adapted for rapid digging, their limbs are short and have strong [[claw]]s. The claws on the hind feet are elongated and curve backwards to enable cleaning and grooming between the spines. Like the Platypus it has a low [[body temperature]] — between 30 and 32 [[Celsius|°C]], but unlike the Platypus, which shows no evidence of [[torpor]] or hibernation, the body temperature of the echidna may fall as low as 5 °C.<ref>Dawson, T.J., Grant, T.R. and Fanning, D. 1979. Standard metabolism of monotremes and the evolution of homeothermy. ''Australian Journal of Zoology'' 27:511–515</ref> The Echidna does not pant or sweat and normally seeks shelter in hot conditions. In autumn and winter the Echidna shows periods of torpor or deep [[hibernation]]. Because of the low body temperature of the Short-beaked Echidna, it becomes sluggish in very hot and very cold weather. Like all monotremes, it has one [[orifice]] for the passage of [[feces|faeces]], [[urine]] and reproductive products, which is known as the [[cloaca]]. The male has internal [[testes]], no external [[scrotum]] and a highly unusual penis with four knobs on the tip. The [[gestation|gestating]] female develops a pouch on its underside, where it raises its young.
[[Image:Echidna, Exmouth.jpg|left|thumb|220px|A Short-beaked Echidna curled into a ball; the snout is visible on the right.]]
The [[muscle|musculature]] of the Short-beaked Echidna has a number of unusual aspects. The [[panniculus carnosus]] is an enormous muscle that is just beneath the skin and covers the entire body. By contraction of various parts of the panniculus carnosus, the Short-beaked Echidna can change shape—the most characteristic shape change is achieved by rolling itself into a ball when threatened, protecting its belly and presenting a defensive array of sharp spines. It has one of the shortest [[spinal cord]]s of any mammal, extending only as far as the [[thorax]].<ref>Hassiotis, M., Paxinos, G., and Ashwell, K.W.S. 2004. Anatomy of the central nervous system of the Australian echidna. ''Proceedings of the Linnean Society of New South Wales'' 125:287–300</ref>
The musculature of the face, jaw and tongue is specialised to allow the Echidna to feed. The tongue of the Short-beaked Echidna is the animal's sole means of catching [[prey]], and can protrude up to 180 mm (8 in) outside the snout. The tongue is sticky because of the presence of [[glycoprotein]]-rich mucous, which both lubricates movement in and out of the snout and helps to catch ants and termites, which adhere to it. Protrusion of the tongue is achieved by contracting the circular muscles that change the shape of the tongue and force it forward, and contraction of two genioglossal muscles attached to the caudal end of the tongue and to the [[mandible]]. The protruded tongue is stiffened by the rapid flow of blood, allowing it to penetrate wood and soil. Retraction requires the contraction of two internal longitudinal muscles, known as the sternoglossi. When the tongue is retracted, the prey is caught on backward-facing [[keratin]]ous "teeth", located along the roof of the buccal cavity, allowing the animal both to capture and grind food.<ref name = "Murray''>Murray, P.F. 1981. A unique jaw mechanism in the echidna, ''Tachglossus aculeatus'' (Monotremata). ''Australian Journal of Zoology'' 29: 1–5</ref> The tongue moves with great speed, and has been measured to move in and out of the snout 100 times a minute.<ref>Doran, G.A. and Baggett, H. 1970. The vascular stiffening mechanism in the tongue of the echidna (''Tachyglossus aculeatus''). ''Anatomical Record'' 167: 197–204</ref>
Numerous [[physiological adaptations]] aid the lifestyle of the Short-beaked Echidna. Because the animal burrows, it can tolerate very high levels of [[carbon dioxide]] in inspired air, and will voluntarily remain in situations where carbon dioxide concentrations are high. Its ear is sensitive to low-[[frequency]] sound, which may be ideal for detecting sounds emitted by termites and ants underground. The leathery snout is covered in mechano- and thermoreceptors, which provide information about the surrounding environment.<ref>Iggo, A., McIntyre, A.K. & Proske, U. (1985). Responses of mechanoreceptors and thermoreceptors in skin of the snout of the echidna ''Tachyglossus aculeatus''. ''Proceedings of the Royal Society of London B'' 223: 261–277</ref> The Short-beaked Echidna has a well-developed [[olfactory system]], which may be used to detect mates and prey. It has a highly sensitive [[optic nerve]], and has been shown to have visual discrimination and [[spatial memory]] comparable to those of a [[rat]].<ref>Burke, D. 2002. Win-shift and win-stay learning in the short-beaked echidna (''Tachyglossus aculeatus''). ''Animal Cognition'' 5:79–84 </ref> The [[brain]] and [[central nervous system]] of the Short-beaked Echidna have been extensively studied for evolutionary comparison with [[Placentalia|placental mammals]]. The Short-beaked Echidna has the largest [[prefrontal cortex]] with respect to body size of any mammal, it shows [[rapid eye movement during sleep]], and its brain has been shown to contain a [[claustrum]] that is similar to placental mammals, linking this structure to their [[Common descent|common ancestor]].<ref>Nicol, S.C., et al. 2000. The echidna manifests typical characteristics of rapid eye movement sleep. ''Neuroscience Letters'' 283:49–52</ref><ref>
Ashwell, K.W.S., Hardman, C.D., and Paxinos, G. 2004. The claustrum is not missing from all monotreme brains. ''Brain Behaviour and Evolution'' 64:223–241 PMID 15319553</ref>
==Ecology and behaviour==
[[Image:French Island Echidna.ogg|thumb|320px|A Short-beaked Echidna building a defensive burrow on [[French Island National Park|French Island]] (video 0:43s)]]
No systematic study of the ecology of the Short-beaked Echidna has been published; however, there have been studies of several aspects of their ecological behaviour. Short-beaked Echidnas live alone and apart from the burrow created for rearing young; they have no fixed shelter or nest site. They do not have a home territory, but range over a wide area. Short-beaked Echidnas are typically active in the daytime; however, they are ill-equipped to deal with heat, because they have no [[sweat gland]]s and do not pant. Therefore, in warm weather they change their pattern of activity, becoming [[crepuscular]] or [[nocturnal]]. They can tolerate cold temperatures, and hibernate during the winter in very cold regions.<ref>Grigg, G.C., Beard, L.A., and Augee, M.L. 1989. Hibernation in a monotreme, the echidna (''Tachyglossus aculeatus''). ''Comparative Biochemistry and Physiology. A, Comparative Physiology'' 92:609–612 PMID 2566425</ref>
Short-beaked Echidnas can live anywhere that has a good supply of food. Short-beaked Echidnas locate food by smell, using sensors in the tip of their snout, and regularly feast on ants and termites. They are powerful diggers, using their clawed front paws to dig out prey and create burrows for shelter. They may rapidly dig themselves into the ground if they cannot find cover when in danger.
In Australia they are most common in forested areas where there are abundant termite-filled fallen logs. In agricultural areas, they are most likely to be found in uncleared scrub; they may be found in grassland, arid areas, and in the outer suburbs of the capital cities. Little is known about their distribution in New Guinea, they have been found in southern New Guinea between Merauke in the west, to the Kelp Welsh River, east of [[Port Moresby]] in the east, where they may be found in open woodland.<ref name="Flannery"/>
===Reproduction===
[[Image:Ameisenigel Unterseite-drawing.jpg|left|180px|thumb|The underside of a female Short-beaked Echidna; the pouch which carries eggs is towards the rear of the abdomen.]]
The solitary Short-beaked Echidna looks for a mate between May and September; the precise timing of the mating season varies with geographic location. Both males and females give off a strong odour during the mating season. During courtship — observed for the first time in 1989 — males locate and pursue females. Trains of up to ten males may follow a single female in a courtship ritual that may last for up to four weeks; the duration of the courtship period varies with location.<ref>Rismiller, P.D., Seymour, R.S., 1991. The echidna. ''Scientific American'' 264, 96–103.</ref> In cooler parts of their range, such as Tasmania, females may mate within a few hours of arousal from hibernation.<ref>Nicol, S.C., Andersen, N.A., and Jones, S.M., 2005. Seasonal variations in reproductive hormones of free-ranging echidnas (''Tachyglossus aculeatus''): interaction between reproduction and hibernation. ''General and Comparative Endocrinology'' 144: 204–210</ref>
Before mating, the male smells the female, paying particular attention to the [[cloaca]]. The male is often observed to roll the female onto her side and then assumes a similar position so that the two animals are abdomen to abdomen. Each side of the bilaterally symmetrical, rosettelike 4 headed penis [similar to reptiles] is used alternately, with the other half being shut down between ejaculations. Sperm bundles of ~100 each, appears to confer increased sperm motility, which may provide the potential for sperm competition between males.<ref>Johnston SD, Smith B, Pyne M, Stenzel D, Holt WV. One-Sided Ejaculation of Echidna Sperm Bundles (''Tachyglossus aculeatus''). ''Am. Nat.'' 2007. Vol. 170, p. E000. [AN:42629 DOI:10.1086/522847]</ref> Each mating results in the production of a single egg, and females are known to mate only once during the breeding season; each mating is successful.<ref>Rismiller, P.D. and McKelvey, M.W. 2000. Frequency of breeding recruitment in the Short-beaked Echidna, ''Tachyglossus aculeatus''. ''Journal of Mammalogy'' 81:1–17</ref>
Fertilisation occurs in the [[oviduct]]. [[Gestation]] takes between 21 and 28 days, during which time the female constructs a nursery burrow. Following the gestation period, a single rubbery-skinned egg between 13 and 17 millimetres in diameter is laid directly into a small, backward-facing pouch that has developed on her abdomen. Ten days after it is laid, the egg hatches within the pouch. The embryo develops an "egg tooth" during incubation, which it uses to tear open the egg; the tooth disappears soon after hatching.
Hatchlings are about 1.5 cm long and weigh between 0.3 and 0.4 [[gram]]s.<ref name = "Rismiller2003">Rismiller, P.D. and McKelvey, M.W. 2003. Body mass, age and sexual maturity in short-beaked echidnas, ''Tachyglossus aculeatus''. ''Comparative Biochemistry and Physiology. A, Molecular & Integrative Physiology'' 136:851–865</ref> After hatching, young Echidnas are known as ''puggles''. Hatchlings attach themselves to their mothers' milk areolae, a specialised patch on the skin that secretes milk (monotremes lack nipples). The way in which puggles imbibe the milk is not yet known, but they have been observed ingesting large amounts during each feeding period, since mothers may leave them unattended in the burrow for between five and ten days. The principal components of the milk are fucosyllactose and saialyllactose; it is high in iron content, giving it a pink colour. Juveniles are eventually ejected from the pouch at around two to three months of age, because of the continuing growth in the length of their spines. Suckling gradually decreases until juveniles are weaned at about six months of age. The duration of lactation is about 200 days, and the young leave the burrow between 180 and 240 days.
The age of sexual maturity is uncertain, but may be four to five years. A twelve-year field study, published in 2003, found that the Short-beaked Echidna reached sexual maturity between five and 12 years of age, and that the frequency of reproduction varies from once every two years to once every six years.<ref name = "Rismiller2003">Rismiller, P.D. and McKelvey, M.W. 2003. Body mass, age and sexual maturity in short-beaked echidnas, ''Tachyglossus aculeatus''. ''Comparative Biochemistry and Physiology. A, Molecular & Integrative Physiology'' 136:851–865</ref> The Short-beaked Echidna can live as long as 45 years in the wild.
Similar to its fellow monotreme the Platypus, the Short-beaked Echidna has an unusual system of sex chromosomes, resulting in males having one fewer chromosomes than females. Male individuals appear to be XYXYXYXYX, with the final X unpaired, while females are XXXXXXXXXX. Weak homology between chromosomes results in meiotic pairing that yields only two possible genotypes of sperm, XXXXX or YYYY, thus preserving this complex system.<ref name = "Rensetal2007">{{cite journal | author = Rens, W., et al | title = The multiple sex chromosomes of platypus and echidna are not completely identical and several share homology with the avian Z | journla = Genome Biology | volume = 8 | pages = R243}}</ref>
==Conservation status==
[[Image:Echidna - melbourne zoo.jpg|right|220px|thumb|A Short-beaked Echidna on the move]]
The Short-beaked Echidna is common throughout most of temperate Australia and lowland New Guinea, and is not listed as endangered. In Australia, the number of Short-beaked Echidnas has been less affected by land clearing than have some other species, since Short-beaked Echidnas do not require a specialised habitat beyond a good supply of ants and termites. Despite their spines, they are preyed on by [[birds of prey|birds]], the [[Tasmanian Devil]], cats, foxes and dogs. They were eaten by [[indigenous Australians]] and the early European settlers of Australia. The most common threats to the animal in Australia are motorised vehicles and habitat destruction, which have led to localised extinction. Infection with the introduced parasite ''[[Spirometra erinaceieuropaei]]'' is fatal for the Echidna. The Wildlife Preservation Society of Queensland runs an Australia-wide survey called ''Echidna Watch'' to monitor the species in Australia.
Captive breeding is difficult, partly due to the relatively infrequent breeding cycle. Only five zoos have managed to breed a captive Short-beaked Echidna, but no captive-bred young have survived to maturity. This has conservation implications for the endangered species of echidna from the genus ''Zaglossus'', and to a lesser extent for the Short-beaked Echidna.
==Cultural references==
[[Image:Australian Five Cents.jpg|thumb|right|260px|Short-beaked Echidna on the Australian five-cent piece (on the left)]]
Short-beaked Echidnas feature in the [[animism|animistic]] culture of Indigenous Australians, including their [[Australian Aboriginal art|visual arts]] and stories. The species was a totem for some groups, including the [[Noongar|Noongar people]] from Western Australia. Many groups have myths about the animal; one myth explains that it was created when a group of hungry, young men went hunting at night and stumbled across a [[wombat]]. They threw spears at the wombat, but lost sight of it in the darkness. The wombat adapted the spears as its own defense and turned into an Echidna.<ref>Robinson, R. 1966. ''Aboriginal Myths and Legends''. Sun Books, Melbourne</ref> Another story tells of a greedy man that kept food from his tribe; warriors speared him and he crawled away into the bushes, where he was turned into an Echidna, the spears becoming his spines.<!--numerous groups-->
The Short-beaked Echidna is an iconic animal in contemporary Australia, notably appearing on the [[Australian dollar|Australian]] [[Australian currency|five-cent piece]] (the smallest denomination) and on a $200 commemorative coin released in 1992. The Short-beaked Echidna has been included in several postal issues: it was one of four native species to appear on Australian postage stamps in 1974, where it was the 25 cent stamp; it appeared on a 37 cent stamp in 1987, and again in 1992 when it was on the 35 cent stamp. The anthropomorphic Echidna ''Millie'' was a mascot for the [[2000 Summer Olympics#mascot|2000 Summer Olympics]].
==See also==
*[[Fauna of Australia]]
*[[List of monotremes and marsupials]]
==References==
===Cited references===
{{Reflist|2}}
===General references===
<div class="references-small">
*Augee, M and Gooden, B. 1993. ''Echidnas of Australia and New Guinea''. Australian National History Press, Sydney ISBN 978-0-86840-046-4
*Augee, M.L. 1983. R. Strahan Ed. ''The Australian Museum Complete Book of Australian Mammals''. p. 8–9. Angus & Robertson ISBN 0-207-14454-0
*Griffiths, M. 1989. [http://www.deh.gov.au/biodiversity/abrs/publications/fauna-of-australia/pubs/volume1b/15-ind.pdf Tachyglossidae]. Pp. 407–435 in ''Fauna of Australia'' (D. W. Walton and B. J. Richardson, eds.). Mammalia, Canberra, Australian Capital Territory 1B:1–1227.
</div>
==External links==
{{wikispecies|Tachyglossus aculeatus}}
{{commons|Tachyglossidae}}
{{Spoken Wikipedia|Short-beaked_Echidna.ogg|2006-06-26}}
*[http://www.dpiwe.tas.gov.au/inter.nsf/WebPages/BHAN-5357K5?open Tasmanian Department of Water, Primary Industries and Environment - Short-beaked Echidna]
*[http://www.abc.net.au/science/scribblygum/June2000/default.htm Echidna Love Trains], article with sound clips and images of puggles
*[http://www.wildlife.org.au/wildlife.php?page=w-echidna.html Wildlife Preservation Society of Queensland - Echidna Watch]
*[http://www.australianmammals.org.au/Species/Echidna%20Profile.htm Australian Mammal Species Files]
{{featured article}}
{{Monotremata}}
[[Category:Monotremes]]
[[Category:Mammals of New Guinea]]
[[Category:Myrmecophagous mammals|Echidna, Short-beaked]]
[[Category:Mammals of South Australia]]
[[Category:Mammals of Tasmania]]
[[Category:Mammals of Western Australia]]
[[Category:Mammals of the Northern Territory]]
[[Category:Mammals of New South Wales]]
[[Category:Mammals of Victoria]]
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