Spinocerebellar tract
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215844256
2008-05-29T23:15:12Z
Calvin 1998
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Reverted edits by [[Special:Contributions/72.192.28.238|72.192.28.238]] ([[User talk:72.192.28.238|talk]]) to last version by 90.197.24.144
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The '''spinocerebellar tract''' is a set of [[axon|axonal]] fibers originating in the [[spinal cord]] and terminating in the ipsilateral [[cerebellum]]. This tract conveys information to the cerebellum about limb and joint position ([[proprioception]]).
==Origins of proprioceptive information==
Proprioceptive information is obtained by [[Golgi organ|Golgi tendon organs]] and [[Muscle spindle|muscle spindles]].
* [[Golgi organ|Golgi tendon organs]] consist of a fibrous capsule enclosing tendon fasciculi and bare nerve endings that respond to tension in the tendon by causing [[action potential]]s in [[1β afferent neurones]] (relatively large, myelinated, quickly conducting).
* [[Muscle spindle|Muscle spindles]] fibres are complicated systems of tension monitoring within muscles which result in information being carried via [[1α neurones]] (larger and faster than 1β) (from both [[nuclear bag fibres]] and [[nuclear chain fibres]]) and [[II neurones]] (solely from nuclear chain fibres).
All of these neurones are "first order" or "primary", are sensory (and thus have their cell bodies in the [[dorsal root ganglion]]) and pass through [[Rexed laminae]] layers I-VI of the [[dorsal horn]], to form synapses with "second order" or "secondary" neurons in the layer just beneath the dorsal horn (layer VII)
==Subdivisions of the tract==
The tract is divided into:
{| class="wikitable"
| '''Division''' || '''Information''' || '''Limbs'''
|-
| [[Dorsal spinocerebellar tract|dorsal (posterior) spinocerebellar tract]] || Information from [[muscle spindle]]s || hind limbs
|-
| [[Ventral spinocerebellar tract|ventral (anterior) spinocerebellar tract]] || Information from [[Golgi organ|golgi tendon organs]] || hind limbs
|-
| [[spinocuneocerebellar tract]] || Information from [[muscle spindle]]s || forelimbs
|-
| [[rostral spinocerebellar tract]] || Information from [[Golgi organ|golgi tendon organs]] || forelimbs
|}
==Pathway for dorsal and spinocuneocerebellar tracts==
In the dorsal tract, the sensory neurones synapse in an area known as '''[[Clarke's nucleus]]''' or "Clarke's column".
This is a column of relay neurone cell bodies within the medial gray matter within the [[spinal cord]] in layer VII (just beneath the dorsal horn), specifically between T1-L2. These neurones then send axons up the [[spinal cord]] and form synapses in the accessory (lateral) [[cuneate nucleus]], lateral to the cuneate nucleus in the medulla.
Below L2, relevant neurones pass into the [[fasciculus gracilis]] (usually associated with the dorsal column-medial lemniscal system) until L2 where they synapse with Clarke's nucleus (leading to considerable caudal enlargement).
From above T1, neurones enter the [[fasciculus cuneatus]] directly and again synapse with neurones in the accessory cuneate nucleus. This pathway is known as the '''spinocuneocerebellar tract'''.
The neurones in the accessory cuneate nucleus have axons leading to the ipsilateral cerebellum via the caudal cerebellar peduncle.
==Pathway for ventral and rostral tracts==
Some neurones instead form synapses with neurones in layer VII of L4-S3. Most of these fibres cross over to the contralateral lateral funiculus via the [[anterior white commissure]] and proceed up the spinal cord to synapse with neurones in the [[superior cerebellar peduncle]]. The fibres then often cross over again within the cerebellum to end on the ipsilateral side.
The Rostral Tract is similar but is uncrossed and enters the cerebellum through both the superior and [[inferior cerebellar peduncle]]s.
==External links==
* http://www.anatomyatlases.org/MicroscopicAnatomy/Section17/Plate17327.shtml
{{Spinal cord}}
{{Neural tracts}}
[[Category:Central nervous system]]
[[Category:Neuroanatomy]]