Swift
219023
226004616
2008-07-16T12:08:17Z
81.207.118.238
{{otheruses}}
{{Taxobox
| name = Swifts
| image = Apus apus 01.jpg
| image_width = 220px
| image_caption = [[Common Swift]], ''Apus apus''<br/>Note wing shape different from [[swallow]]s
| regnum = [[Animal]]ia
| phylum = [[chordate|Chordata]]
| classis = [[bird|Aves]]
| ordo = [[Apodiformes]]
| familia = '''Apodidae'''
| familia_authority = [[Ernst Hartert|Hartert]], [[1897]]
| subdivision_ranks = Genera
| subdivision =
Nearly 20, see text.
}}
The '''swifts''' are [[bird]]s superficially similar to [[swallow]]s but are actually not closely related to those [[passerine]] species at all; swifts are in the separate order Apodiformes, which they share with the [[hummingbird]]s. The [[treeswift]]s are closely related to the true swifts, but form a separate family, the Hemiprocnidae.
The resemblances between swifts and swallows are due to [[convergent evolution]] reflecting similar life styles based on catching [[insect]]s in flight.
The family scientific name comes from the [[Ancient Greek]] απους, ''apous'', meaning "without feet", since swifts have very short legs and never settle voluntarily on the ground, perching instead on vertical surfaces. The tradition of depicting swifts without feet continued into the Middle Ages, as seen in the heraldic [[martlet]].
==Description==
Swifts are the most aerial of birds and some, like the [[Common Swift]], even sleep and mate on the wing. Larger species, such as [[White-throated Needletail]], are amongst the fastest flyers in the animal kingdom. One group, the [[Swiftlet]]s or Cave Swiftlets have developed a form of [[Animal echolocation|echolocation]] for navigating through dark cave systems where they roost<ref name=EoB/>. One species, ''Aerodramus papuensis'' has recently been discovered to use this navigation at night outside its cave roost also.
Swifts have a worldwide distribution in tropical and temperate areas, but like swallows and martins, the swifts of temperate regions are strongly [[bird migration|migratory]] and winter in the tropics. Some species can survive short periods of cold weather by entering [[torpor]], a state similar to hibernation<ref name=EoB>{{cite book |editor=Forshaw, Joseph|author= Collins, Charles T.|year=1991|title=Encyclopaedia of Animals: Birds|publisher= Merehurst Press|location=London|pages= 134-136|isbn= 1-85391-186-0}}</ref>.
Many swifts have a characteristic shape, with a short forked tail and very long swept-back wings that resemble a crescent or a [[boomerang]]. The flight of some species is characterised by a distinctive "flicking" action quite different from swallows. Swifts range in size from the Pygmy Swiftlet (''Collocalia troglodytes''), which weighs 5.4 g and measures 9 cm (3.7 inches) long, to the [[Purple Needletail]] (''Hirundapus celebensis''), which weighs 184 g (6.5 oz) and measures 25 cm (10 inches) long<ref name=EoB/>.
The nest of many species is glued to a vertical surface with saliva, and the genus ''[[Aerodramus]]'' use only that substance, which is the basis for [[bird's nest soup]]. The eggs hatch after 19 to 23 days, and the young leave the nest after a further six to eight weeks. Both parents assist in raising the young<ref name=EoB/>.
==Systematics and evolution==
Swifts and treeswifts have long been considered to be relatives of the [[hummingbird]]s, a judgement corroborated by the discovery of the [[Jungornithidae]], which were apparently swift-like hummingbird relatives, and of primitive hummingbirds such as ''[[Eurotrochilus]]''. Traditional taxonomies place the hummingbird family (Trochilidae) in the same order as the swifts; the [[Sibley-Ahlquist taxonomy]] places them in a new order, Trochiliformes, which forms a superorder with the swifts and treeswifts (and no other birds).
The taxonomy of the swifts is in general complicated, with genus and species boundaries widely disputed, especially amongst the [[swiftlet]]s. Analysis of behavior and vocalizations is marred by common [[parallel evolution]], while analyses of different [[Morphology (biology)|morphological traits]] and of various [[DNA sequences]] have yielded equivocal and partly contradictory results (Thomassen ''et al.'', 2005).
The Apodiformes diversified during the [[Eocene]], at the end of which the extant families were present; fossil genera are known from all over temperate Europe, between today's [[Denmark]] and [[France]], such as the primitive ''[[Scaniacypselus]]'' (Early - Middle Eocene) and the more modern ''[[Procypseloides]]'' (Late Eocene/Early [[Oligocene]] - Early [[Miocene]])<!-- Geobios36:719 -->. A prehistoric genus sometimes assigned to the swifts, ''[[Primapus]]'' (Early Eocene of England), might also be a more distant ancestor.
===Taxonomic list of Apodidae===
'''Tribe Cypseloidini'''
* Genus ''[[Cypseloides]]'' - includes ''Streptoprocne''
** [[Chestnut-collared Swift]], ''Cypseloides rutilus''
** [[Tepui Swift]], ''Cypseloides phelpsi''
** [[Black Swift]], ''Cypseloides Niger''
** [[White-chested Swift]], ''Cypseloides lemosi''
** [[Rothschild's Swift]], ''Cypseloides rothschildi''
** [[Sooty Swift]], ''Cypseloides fumigatus''
** [[Spot-fronted Swift]], ''Cypseloides cherriei''<!-- Bull. B.O.C. 2004 124:154 -->
** [[White-chinned Swift]], ''Cypseloides cryptus''
** [[White-fronted Swift]], ''Cypseloides storeri''
** [[Great Dusky Swift]], ''Cypseloides senex''
* Genus ''[[Streptoprocne]]''
** [[White-collared Swift]], ''Streptoprocne zonaris''
** [[Biscutate Swift]], ''Streptoprocne biscutata''
** [[White-naped Swift]], ''Streptoprocne semicollaris''
'''Tribe Collocalini''' - [[swiftlet]]s
* Genus ''[[Collocalia]]'' (3-4 species)
* Genus ''[[Aerodramus]]'' (about 25 species, sometimes included in ''Collocalia'')
* Genus ''[[Hydrochous]]'' - Waterfall Swift
* Genus ''[[Schoutedenapus]]'' - African swiftlets (2 species)
'''Tribe Chaeturini''' - needletails
* Genus ''[[Mearnsia]]''
** [[Philippine Needletail]], ''Mearnsia picina''
** [[Papuan Spinetail]], ''Mearnsia novaeguineae''
* Genus ''[[Zoonavena]]''
** [[Malagasy Spinetail]], ''Zoonavena grandidieri''
** [[Sao Tome Spinetail]], ''Zoonavena thomensis''
** [[White-rumped Needletail]], ''Zoonavena sylvatica''
* Genus ''[[Telacanthura]]''
** [[Mottled Spinetail]], ''Telacanthura ussheri''
** [[Black Spinetail]], ''Telacanthura melanopygia''
* Genus ''[[Rhaphidura]]''
** [[Silver-rumped Needletail]], ''Rhaphidura leucopygialis''
** [[Sabine's Spinetail]], ''Rhaphidura sabini''
* Genus ''[[Neafrapus]]''
** [[Cassin's Spinetail]], ''Neafrapus cassini''
** [[Bat-like Spinetail]], ''Neafrapus boehmi''
* Genus ''[[Hirundapus]]''
** [[White-throated Needletail]], ''Hirundapus caudacutus''
** [[Silver-backed Needletail]], ''Hirundapus cochinchinensis''
** [[Brown-backed Needletail]], ''Hirundapus giganteus''
** [[Purple Needletail]], ''Hirundapus celebensis''
* Genus ''[[Chaetura]]'' (9 species)
'''Tribe Apodini''' - typical swifts
* Genus ''[[Aeronautes]]''
** [[White-throated Swift]], ''Aeronautes saxatalis''
** [[White-tipped Swift]], ''Aeronautes montivagus''
** [[Andean Swift]], ''Aeronautes andecolus''
* Genus ''[[Tachornis]]''
** ''Tachornis uranoceles'' ([[fossil]]; Late Pleistocene of Puerto Rico)<!-- Auk 99:230 -->
** [[Antillean Palm Swift]], ''Tachornis phoenicobia''
** [[Pygmy Swift]], ''Tachornis furcata''
** [[Fork-tailed Palm Swift]], ''Tachornis squamata''
* Genus ''[[Panyptila]]''
** [[Lesser Swallow-tailed Swift]], ''Panyptila cayennensis''
** [[Great Swallow-tailed Swift]], ''Panyptila sanctihieronymi''
* Genus ''[[Cypsiurus]]''
** [[Asian Palm Swift]], ''Cypsiurus balasiensis''
** [[African Palm Swift]], ''Cypsiurus parvus''
* Genus ''[[Apus (genus)|Apus]]'' (some 17 species)
==References==
{{reflist}}
{{commonscat|Apodidae}}
* '''Chantler''', Phil & '''Driessens''', Gerald (2000): ''Swifts : a guide to the swifts and treeswifts of the world''. Pica Press, Mountfield, East Sussex. ISBN 1-873403-83-6
* '''Thomassen''', Henri A.; Tex, Robert-Jan; de Bakker, Merijn A.G. & Povel, G. David E. (2005): Phylogenetic relationships amongst swifts and swiftlets: A multi locus approach. ''Molecular Phylogenetics and Evolution'' '''37'''(1): 264-277. {{DOI|10.1016/j.ympev.2005.05.010}} (HTML abstract)
==External links==
* [http://commonswift.org/common_swift.html CommonSwift.org] -- Comprehensive website in several languages
*[http://www.xeno-canto.org/browse.php?query=apodidae Swift sounds] on xeno-canto.org
[[Category:Apodidae|*]]
[[Category:swifts|*]]
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