Tongue
55999
225816163
2008-07-15T15:17:29Z
J.delanoy
2372780
Reverted edits by [[Special:Contributions/90.201.232.110|90.201.232.110]] to last version by 89.250.190.68 (using [[WP:HG|Huggle]])
{{otheruses}}{{Refimprove|date=June 2008}}
{{Infobox Anatomy |
Name = Tongue |
Latin = lingua |
GraySubject = 242 |
GrayPage = 1125 |
Image = Tongue.agr.jpg |
Caption = A [[human]] tongue |
Image2 = |
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Precursor = [[pharyngeal arches]], [[lateral lingual swelling]], [[tuberculum impar]]<ref>{{EmbryologyUNC|hednk|024}}</ref> | ccjvjkdhfgrghg
Artery = [[lingual artery|lingual]], [[Tonsillar branch of the facial artery|tonsillar branch]], [[ascending pharyngeal artery|scending pharyngeal]] |
Vein = [[lingual veins|lingual]] |
Nerve = [[lingual nerve]] |
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MeshNumber = A03.556.500.885 |
DorlandsPre = l_11 |
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}}
The '''tongue''' is the large bundle of [[skeletal muscle]]s on the floor of the [[mouth]] that manipulates [[food]] for [[chewing|chewing and swallowing]] (deglutition). It is the primary organ of [[taste]]. Much of the surface of the tongue is covered in [[taste bud]]s. The tongue, with its wide variety of possible movements, assists in forming the sounds of [[Speech communication|speech]]. It is sensitive and kept moist by [[saliva]], and is richly supplied with nerves and blood vessels to help it move.<ref>{{cite book
| last = Maton
| first = Anthea
| authorlink =
| coauthors = Jean Hopkins, Charles William McLaughlin, Susan Johnson, Maryanna Quon Warner, David LaHart, Jill D. Wright
| title = Human Biology and Health
| publisher = Prentice Hall
| date = 1993
| location = Englewood Cliffs, New Jersey, USA
| pages =
| url =
| doi =
| id =
| isbn = 0-13-981176-1}}</ref>
==Etymology==
The word '''tongue''' can be used as a [[metonymy]] for ''language'', as in the phrase '''[[mother tongue]]'''. In fact, [[Albanian language|Albanian]] (''gjuha''), [[Catalan language|Catalan]] (''llengua''), [[Portuguese language|Portuguese]] (''língua''), [[French language|French]] (''langue''), [[Maltese language|Maltese]], (''ilsien''), [[Arabic]] (لسان ''lisān''), [[Romanian language|Romanian]] (''limba''), [[Russian language|Russian]] (язык ''yazyk''), [[Bulgarian language|Bulgarian]] (''ezik''), [[Persian language|Persian]] (''zabaan''), [[Greek language|Greek]] (γλώσσα), [[Spanish language|Spanish]] (''lengua''), [[Polish language|Polish]], [[Slovak language|Slovak]], [[Czech language|Czech]], [[Slovene language|Slovene]], [[Bosnian language|Bosnian]], [[Serbian language|Serbian]], [[Croatian language|Croatian]] (''jezik''), [[Armenian language|Armenian]] (լեզու), [[Finnish language|Finnish]] (''kieli''), [[Estonian language|Estonian]] (''keel''), [[Irish language|Irish]], [[Italian language|Italian]] (''lingua''), [[Latin]] (''lingua''), [[Urdu]] (''zabaan''), [[Aramaic language|Aramaic]] (ܠܫܢܐ/לשנא ''lišānā''), [[Hungarian language|Hungarian]] (''nyelv''), [[Hebrew language|Hebrew]] (לָשׁוֹן ''lashon''), [[Turkish language|Turkish]] (''dil''), and [[Danish language|Danish]] (''tunge''), all have the same word for "tongue" and "[[language]]". [[Webster's Dictionary]] initially spelt the word ''Tung''.{{Fact|date=July 2008}}
===Figures of speech===
A common temporary failure in word [[Recollection|retrieval]] from [[memory]] is referred to as the ''[[Tip of the tongue|tip-of-the-tongue]]'' [[phenomenon]]. The expression ''[[Tongue-in-cheek|tongue in cheek]]'' refers to a statement that is not to be taken entirely seriously; something said or done with subtle ironic humour. "Tongue twisted" is a term used to described being unable to pronounce a word or phrase correctly. A ''[[tongue twister]]'' is a phrase made specifically to be very difficult to pronounce. "Tongue-tied" means being unable to say what you want to due to confusion or restriction. The phrase "cat got your tongue" refers to when a person is speechless.
==Anatomy==
=== Structure ===
[[Image:Gray1014.png|right|thumb|Drawing of an anterior view of the tongue and oral cavity, with cheeks removed for clarity.]]
[[Image:Gray1019.png|thumb|Lateral view of the tongue, with extrinsic muscles highlighted.]]
The tongue is made mainly of [[skeletal muscle]]. The tongue extends much further than is commonly perceived, past the posterior border of the mouth and into the oropharynx.
The dorsum (upper surface) of the tongue can be divided into two parts:
* an '''oral part''' (anterior two-thirds of the tongue) that lies mostly in the mouth
* a '''pharyngeal part''' (posterior third of the tongue), which faces backward to the [[oropharynx]]
The two parts are separated by a V-shaped groove, which marks the [[Terminal sulcus (tongue)|sulcus terminalis]] (or ''terminal sulcus'').
Since the tongue contains no bony supports for the muscles, the tongue is an example of a [[muscular hydrostat]], similar in concept to an [[octopus]] arm. Instead of bony attachments, the '''extrinsic muscles of the tongue''' anchor the tongue firmly to surrounding bones and prevent the mythical possibility of 'swallowing' the tongue.
Other divisions of the tongue, are based on the area of the tongue:
{| class="wikitable"
|-
! normal name
! anatomical name
! adjective
|-
| tongue tip
| apex
| apical
|-
| tongue blade
| lamina
| laminal
|-
| tongue dorsum
| dorsum (back)
| dorsal
|-
| tongue root
| radix
| radical
|-
| tongue body
| corpus
| corporeal
|}
===Muscles of the tongue===
The intrinsic muscles lie entirely within the tongue, while the extrinsic muscles attach the tongue to other structures.
[[Image:que lengua.jpg|thumb|3/4 view of a 6.5 cm human tongue.]]
The extrinsic muscles reposition the tongue, while the intrinsic muscles alter the shape of the tongue for talking and swallowing.
====Extrinsic muscles====
Extrinsic muscles of the tongue by definition originate from structures outside the tongue and insert into the tongue. The four paired extrinsic muscles protrude, retract, depress, and elevate the tongue:
{| class="wikitable"
| '''Muscle''' || '''From''' || '''Nerve''' || '''Function'''
|-
| ''[[Genioglossus muscle]]'' || [[mandible]] || [[hypoglossal nerve]] || protrudes the tongue as well as depressing its center.
|-
| ''[[Hyoglossus muscle]]'' || [[hyoid bone]] || [[hypoglossal nerve]] || depresses the tongue.
|-
| ''[[Styloglossus muscle]]'' || [[styloid process]] || [[hypoglossal nerve]] || elevates and retracts the tongue.
|-
| ''[[Palatoglossus muscle]]'' || [[palatine aponeurosis]] || [[pharyngeal branch of vagus nerve]] || depresses the [[soft palate]], moves the ''palatoglossal fold'' towards the midline, and elevates the back of the tongue.
|}
====Intrinsic muscles====
[[Image:Gray1020.png|thumb|Coronal section of tongue, showing intrinsic muscles]]
Four paired intrinsic muscles of the tongue originate and insert within the tongue, running along its length. These muscles alter the shape of the tongue by: lengthening and shortening it, curling and uncurling its apex and edges, and flattening and rounding its surface.<ref>Drake, R. et al. ''Gray's Anatomy for Students,'' Elsevier, 2005. ISBN 0-443-06612-4</ref>
*The ''[[superior longitudinal muscle]]'' runs along the superior surface of the tongue under the mucous membrane, and elevates, assists in retraction of, or deviates the tip of the tongue. It originates near the [[epiglottis]], the [[hyoid bone]], from the median fibrous septum.
*The ''[[inferior longitudinal muscle]]'' lines the sides of the tongue, and is joined to the styloglossus muscle.
*The ''[[verticalis muscle]]'' is located in the middle of the tongue, and joins the superior and inferior longitudinal muscles.
*The ''[[transversus muscle]]'' divides the tongue at the middle, and is attached to the [[mucous membrane]]s that run along the sides.
The tongue is often cited as the "[[Muscle#The "strongest" human muscle|strongest muscle in the body]]," a claim that does not correspond to any conventional definition of strength.
===Papillae and taste buds===
{{See also|Taste bud}}
The oral part of the tongue is covered with small bumpy projections called [[wiktionary:papillae|papillae]]. There are four types of papillae:
* [[filiform]] (thread-shape)
* [[fungiform papilla|fungiform]] (mushroom-shape)
* [[circumvallate papilla|circumvallate]] (ringed-circle)
* [[foliate papilla|foliate]] (leaf-shape)
All papillae except the filiform have [[taste bud]]s on their surface.
[[Image:Tongue tastebuds.jpg|left|thumb|Close-up view of a tongue with visible [[fungiform papillae]] (large bumps) scattered among [[filiform papillae]] (small bumps).]]
The circumvallate are the largest of the papillae. There are 8 to 14 circumvallate papillae arranged in a V-shape in front of the sulcus terminalis, creating a border between the oral and pharyngeal parts of the tongue.
There are no lingual papillae on the underside of the tongue. It is covered with a smooth [[mucous membrane]], with a fold (the [[lingual frenulum]]) in the center. If the lingual frenulum is too taut or too far forward, it can impede motion of the tongue, a condition called [[Tongue-tie (Ankyloglossia)]].
The upper side of the posterior tongue (pharyngeal part) has no visible taste buds, but it is bumpy because of the lymphatic nodules lying underneath. These follicles are known as the [[lingual tonsil]].
The human tongue can detect five basic taste components: [[sweet]], [[sour]], [[salty]], [[Basic taste#Bitterness|bitter]] and [[savory]]{{Fact|date=January 2008}}. The sense of taste is referred to as a ''gustatory'' sense. Contrary to the popular myth and generations of schoolbooks, there are no distinct regions for tasting different tastes. This myth arose because [[Edwin G. Boring]] replotted data from one of [[Wundt]]'s students (Hanig) without labeling the axes, leading some to misinterpret the graph as all or nothing response.<ref>Bartoshuk, L.M. (1989). Taste: Robust across the Age Span? Annals of the New York Academy of Sciences 561, pp. 65-75.</ref> The common conception of ''taste'' has a significant contribution from [[olfaction]].
=== Innervation of the tongue ===
''Motor innervation'' of the tongue is complex and involves several cranial nerves. All the muscles of the tongue are innervated by the [[hypoglossal nerve]] (cranial nerve XII) with one exception: the palatoglossal muscle is innervated by the [[pharyngeal branch of vagus nerve]] (cranial nerve X).
''Sensory innervation'' of the tongue is different for taste sensation and general sensation.
* For the ''[[anterior tongue|anterior two-thirds of the tongue]]'', general sensations and taste sensations are carried via different nerves.
** Somatic sensations travel from the tongue via the [[lingual nerve]], a major branch of the [[mandibular nerve]] (itself a branch of the [[trigeminal nerve]]). This nerve also carries general sensation from areas of the oral mucosa and [[gingiva]] of the lower teeth.
** Taste sensation is carried to the facial nerve via the [[chorda tympani]]. The chorda tympani also carries parasympathetic fibers from the facial nerve to the [[submandibular ganglion]].
* The ''[[posterior tongue|posterior one-third of the tounge]]'' has a more simple innervation, as both taste and general sensations are carried by the [[glossopharyngeal nerve]].
=== Vasculature of the tongue ===
[[Image:P6231133.JPG|right|thumb|250px|The underside of a human tongue]]
The tongue receives its blood supply primarily from the [[lingual artery]], a branch of the [[external carotid artery]]. The floor of the mouth also receives its blood supply from the lingual artery.
There is also secondary blood supply to the tongue from the [[tonsillar branch]] of the [[facial artery]] and the [[ascending pharyngeal artery]].
==Length==
The average length of the tongue from the oropharynx to the tip is 10 cm (4 in).<ref>{{cite book |year=1997 |author=Robin Kerrod |title=MacMillan's Encyclopedia of Science |publisher=Macmillan Publishing Company, Inc. |volume=6 |isbn=0028645588}}</ref> [[Stephen Taylor]] holds the world record for the [[Largest body part|world's longest tongue]]. It measures 9.5 centimeters from the tip to the center of his closed top lip. Annika Irmler holds the record for longest female tongue, at 7 centimeters.<ref>{{cite web |url=http://news.bbc.co.uk/cbbcnews/hi/world/newsid_1646000/1646912.stm |title='I've got the world's longest tongue' |publisher=CBBC Newsround |date=2001-11-09 |accessed=2007-04-24}}</ref>
== Use of tongue in pharmacy==
The [[sublingual]] region underneath the front of the tongue is a location where the [[oral mucosa]] is very thin, and underlain by a plexus of veins. This is an ideal location for introducing certain medications to the body. The sublingual route takes advantage of the highly [[Blood vessel|vascular]] quality of the oral cavity, and allows for the speedy application of medication into the cardiovascular system, bypassing the gastrointestinal tract. This is the only convenient and efficacious route of administration of [[nitroglycerin]] to a patient suffering [[angina pectoris]], chest pain. If the tablet is swallowed, the medication is completely neutralized by the detoxification process of the [[liver]].{{Fact|date=May 2007}}
== Secondary uses ==
In addition to eating and human vocalization, the human tongue has many secondary uses. These include certain forms of kissing known as "tongue kissing" or sometimes "[[french kiss]]ing" in which the tongue plays a primary role. Generally, use of the tongue (such as licking), or interaction between tongues, appears to be a common gesture of affection, not just in humans but throughout the animal kingdom, and particularly in [[mammal]]s.
The tongue also has a distinct use in both male and female forms of [[oral sex]], and is typically used to a great extent in [[foreplay]] and traditional sexual intercourse as well. Because of its use in both the phenomenon of human sexual interactions, the tongue sometimes is associated with a sensual or erotic connotation. In art the human tongue is often depicted as a seductive instrument, similar to the status of the [[lips]].
[[Image:Tongue ring.JPG|thumb|left|A [[tongue piercing]]]]
The tongue is also one of the more common parts of the human anatomy to be subject to [[piercing]] and [[body modification]], a phenomenon that is sometimes associated with certain subcultures or demographics. [[Tongue piercing]] has appeared historically in many ancient cultures, and is an increasingly popular trend in the West today, particularly in youth culture. Pop culture references to tongue piercings are common as well.
Showing tongue (tongue out) is an international emotional gesture used primarily by children, or by adults behaving (deliberately or not) in a childish manner.
The human tongue also plays a valuable role in other acts, such as for blowing bubbles with [[bubble gum]] and [[whistling]].
Injury to the tongue is often very painful. The muscle is vulnerable to various [[cancer]]s.
==Non-human tongues==
[[Image:Okapitongue.jpg|right|thumbnail|An [[okapi]] using its tongue to [[scratch]] an [[itch]]]]
Most multi-cellular animals, that is, members of the subkingdom [[Metazoa]], have tongues or similar organs.
In animals such as dogs and cats the tongue is often used to clean the fur and body. Rough textures of the tongues of these species helps them to use their tongues to remove oils and parasites by licking themselves and each other. Aside from daily uses for eating and drinking, a dog's tongue acts as a heat regulator. As a dog increases its exercise the tongue will increase in size due to greater blood flow. The tongue hangs out of the dog's mouth and the moisture on the tongue will cool down further cooling down the bloodflow.[http://www.doctordog.com/drdognewsletter/tongue.html][http://www.springerlink.com/content/n3u34u4220384846/]
Some animals have [[prehensile]] tongues. For example, [[chameleon]]s, [[frog]]s, [[salamander]]s and some species of [[fish]] use their tongues to catch prey. Many insects have a type of tongue called a ''[[proboscis]]'' that is used for the same purpose or, in the case of [[butterflies]], to drink [[nectar]] [http://magazine.audubon.org/backyard/backyard.html]. The corresponding organ in [[ant]]s is called the ''[[hypopharynx]]'' [http://jlibsch.web.wesleyan.edu/Ant/Morphology/Head.html]. [[Mollusc]]s have a rough tongue called a ''[[radula]]'' [http://www.qi.com/talk/viewtopic.php?t=917&start=9&sid=a89c9e116f8a414bbfe36cbf6413bc90], which they use to grind food.
== Tongue rolling ==
[[Image:Rolled tongue flikr.jpg|thumb|right|Rolled Tongue]]
Tongue rolling is the act of rolling the tongue axially into a tube shape. The ability to roll the tongue has been generally believed to depend on [[genetic inheritance]]. Tongue rolling was believed to be a [[dominant]] trait with simple [[Mendelian genetics|Mendelian inheritance]], and is still commonly used as an example in high school and introductory [[biology]] courses. It provided a simple experiment to demonstrate inheritance.
There is little laboratory evidence, though, for the common belief that tongue rolling is inheritable and dominant. A 1975 [[twin study]] found that identical twins (who share all of their [[genes]]) were no more likely than fraternal twins (who share an average of half) to both have the same [[phenotype]] for tongue rolling.<ref>[http://www.discovery.com/area/skinnyon/skinnyon970226/skinny1.html Discovery Online, The Skinny On... Tongue Rolling<!-- Bot generated title -->]</ref><ref>[http://www.ncbi.nlm.nih.gov/entrez/dispomim.cgi?id=189300 Omim - Tongue Curling, Folding, Or Rolling<!-- Bot generated title -->]</ref>
==Tongue as a food==
The tongues of some animals are consumed and sometimes considered delicacies. In America and the United Kingdom, cow tongues are among the more common. Hot tongue sandwiches are frequently found on menus in [[Kosher]] [[delicatessen]]s in America. In the United Kingdom tongue can often be found at the local grocer, where it is often sold in reformed slices of meat after being ground up and set in gelatine. [[Taco|Taco de lengua]] (''lengua'' being Spanish for tongue) is a taco filled with [[beef tongue]], and is especially popular in Mexican cuisine. Tongue can also be prepared as [[birria]]. [[Duck]] tongues are sometimes employed in [[Szechuan cuisine|Szechuan]] dishes, while [[Lamb (food)|lamb]]'s tongue is occasionally employed in Continental and contemporary American cooking. Fried [[cod]] tongue is a relatively common part of fish meals in [[Norway]] and [[Newfoundland and Labrador|Newfoundland]].
Tongues are also used in sausage making. Historically, buffalo tongue was once considered an especially exquisite dish, and is one of the reasons for the [[American Bison]] being hunted by humans to the point of near extinction.{{Fact|date=July 2007}}
==See also==
* [[Electronic tongue]]
* [[Language]]
* [[List of Mendelian traits in humans]]
* [[Saliva]]
* [[Taste bud|Taste buds on the tongue]]
* [[Tip of the tongue|Tip-of-the-tongue phenomenon]]
* [[Tongue splitting]]
* [[Tongue cleaner]]
* [[Tongue piercing]]
* [[Tongue-twister]]
* [[Vocal tract]]
* [[Tongue disease]]
==References==
{{reflist}}
==External links==
{{commonscat|tongue}}
{{wiktionary}}
* [http://www.umanitoba.ca/faculties/arts/linguistics/russell/138/sec1/anatomy.htm University of Manitoba, Anatomy of the Vocal Tract]
{{Gustatory system}}
{{human anatomical features}}
{{Head and neck general}}
[[Category:Sensory organs]]
[[Category:Gustatory system]]
[[Category:Mouth]]
[[Category:Digestive system]]
[[ar:لسان]]
[[arc:ܠܫܢܐ (ܐܘܪܓܢܘܢ)]]
[[ast:Llingua (muérganu)]]
[[bs:Jezik (anatomija)]]
[[bg:Език (биология)]]
[[ca:Llengua (múscul)]]
[[cs:Jazyk (orgán)]]
[[cy:Tafod]]
[[da:Tunge]]
[[de:Zunge]]
[[el:Γλώσσα (ανατομία)]]
[[es:Lengua (anatomía)]]
[[eo:Lango (anatomio)]]
[[eu:Mihi]]
[[fa:زبان (کالبدشناسی)]]
[[fr:Langue (anatomie)]]
[[ga:Teanga (anatamaíocht)]]
[[gd:Teanga]]
[[ko:혀]]
[[hr:Jezik (anatomija)]]
[[id:Lidah]]
[[ia:Lingua (anatomia)]]
[[it:Lingua (anatomia)]]
[[he:לשון (איבר)]]
[[pam:Dila]]
[[ka:ენა (ორგანო)]]
[[ku:Ziman (organ)]]
[[la:Lingua (membrum)]]
[[lt:Liežuvis]]
[[ln:Lolému]]
[[hu:Nyelv (testrész)]]
[[mk:Јазик (анатомски орган)]]
[[ml:നാവ്]]
[[ms:Lidah]]
[[nl:Tong (anatomie)]]
[[ja:舌]]
[[no:Tunge]]
[[nn:Tunge]]
[[pag:Dila]]
[[pl:Język (anatomia)]]
[[ro:Limbă (anatomie)]]
[[rmy:Chhib (korposki)]]
[[qu:Qallu]]
[[ru:Язык (анатомия)]]
[[simple:Tongue]]
[[sk:Jazyk (orgán)]]
[[fi:Kieli (anatomia)]]
[[sv:Tunga]]
[[ta:நாக்கு]]
[[te:నాలుక]]
[[th:ลิ้น]]
[[vi:Lưỡi]]
[[tg:Забон (андом)]]
[[tr:Dil (organ)]]
[[uk:Язик]]
[[fiu-vro:Kiil (anatoomia)]]
[[yi:צונג]]
[[zh-yue:脷]]
[[diq:Zıwan (organ)]]
[[zh:舌]]